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2005 Lexus IS300
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input speed sensor usually will fuck with shift points and speedometer, but I've never seen one cause a no run condition but its possible. I have worked on at least 15 auto jz cars and never seen auto cooler lines end there. all radiators come with the ports for cooler lines unless buying one specifically for a manual (dont even think those are sold anymore unless going koyo or mishimoto). When in manual mode you are only telling the car what maximum gear it can shift to but it will auto down shift or upshift if needed.

I would start by replacing the speed sensor.

note that my experience is with lhd usdm is300/gs300s so if you're elsewhere this comment might be irrelevant
 
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